2021 in paleomalacology

From Wikipedia, the free encyclopedia
List of years in paleomalacology
In science
2018
2019
2020
2021
2022
2023
2024
In paleontology
2018
2019
2020
2021
2022
2023
2024
In paleomammalogy
2018
2019
2020
2021
2022
2023
2024
In paleobotany
2018
2019
2020
2021
2022
2023
2024
In arthropod paleontology
2018
2019
2020
2021
2022
2023
2024
In paleoentomology
2018
2019
2020
2021
2022
2023
2024
In paleoichthyology
2018
2019
2020
2021
2022
2023
2024
In reptile paleontology
2018
2019
2020
2021
2022
2023
2024
In archosaur paleontology
2018
2019
2020
2021
2022
2023
2024

This list of fossil molluscs described in 2021 is a list of new taxa of fossil molluscs that were described during the year 2021, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2021.

Ammonites[edit]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Abichites infirmis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Abichites ovalis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Agathiceras iranicum[2]

Sp. nov

Valid

Korn & Ghaderi in Korn et al.

Carboniferous-Permian boundary

 Iran

Alienisphinctes[3]

Gen. et sp. nov

Valid

Énay & Mangold

Late Jurassic (Kimmeridgian)

Jubaila Formation

 Saudi Arabia

A member of the family Ataxioceratidae. The type species is A. cavelieri.

Amaltheus orientalis[4]

Sp. nov

Valid

Nakada et al.

Early Jurassic (Pliensbachian)

Kuruma Group

 Japan

Arasella falcata[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

An ammonite of uncertain placement, probably a member of the family Xenodiscidae.

Arcestes lawsi[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Arnioceras ritterbushi[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

Arnioceras sparsum[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

Artyomites[7]

Gen. et sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Palaeophyllitidae. Genus includes new species A. antiquus.

Aulatornoceras frenklerae[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Aulatornoceras steinhauseni[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Aulatornoceras ventrosulcatum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Boucaulticeras hawthornensis[6]

Sp. nov

Valid

Taylor & Guex

Early Jurassic (Sinemurian)

Sunrise Formation

 United States
( Nevada)

A member of the family Schlotheimiidae.

Bullatimorphites szontaghi[9]

Sp. nov

Valid

Galácz

Caloceras avus[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Catrisimoceras[11]

Gen. et comb. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and possibly to the subfamily Lytogyroceratinae. The type species is "Simoceras" catrianum Zittel (1870).

Cheltonia howarthi[12]

Sp. nov

Valid

Simms & Edmunds

Early Jurassic

 United Kingdom

Cibolaites petraschecki[13]

Sp. nov

Valid

Wilmsen et al.

Late Cretaceous (Turonian)

Brießnitz Formation

 Germany

Clivotirolites[1]

Gen. et 2 sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is C. decoratus; genus also includes C. petilus.

Crassotornoceras hetzeneggeri[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Crassotornoceras nudum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Crioceratites claveli[14]

Sp. nov

In press

Bert et al.

Early Cretaceous (Hauterivian)

Vocontian Basin

 France

Didymoceras morozumii[15]

Sp. nov

Valid

Misaki & Tsujino

Late Cretaceous (Campanian)

Hiketa Formation

 Japan

Dimorphodiscus[16]

Gen. et sp. et comb. nov

Valid

Vermeulen et al.

Early Cretaceous (Barremian)

 Algeria
Crimea
 France

A member of the family Holcodiscidae. The type species is D. lerisae; genus also includes D. uhligi (Karakash, 1907), D. sayni (Joleaud, 1912) and "Metahoplites" postnodosum Vermeulen et al. (2017).

Discoscaphites mullinaxorum[17]

Sp. nov

Valid

Witts et al.

Late Cretaceous (Maastrichtian)

Corsicana Formation

 United States
( Texas)

Domanikoceras martenbergense[18]

Sp. nov

Valid

Korn

Devonian (Frasnian)

 Germany

A member of the family Tornoceratidae.

Dzhulfites brevisellatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Eoasianites baghevangensis[2]

Sp. nov

Valid

Korn & Ghaderi in Korn et al.

Carboniferous-Permian boundary

 Iran

Eoermoceras[3]

Gen. et 2 sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Dhruma Formation

 Saudi Arabia

A member of the family Stephanoceratidae. The type species is E. glabrum; genus also includes new species E. angulatidomus.

Eolytoceras rohrbachensis[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Esfahanites[1]

Gen. et sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is E. armatus.

Eutomoceras paladeji[19]

Sp. nov

Valid

Tongtherm, Nabhitabhata & Tuanapaya

Triassic

Chaiburi Formation

 Thailand

A member of the family Ceratitidae.

Eutomoceras wantanae[19]

Sp. nov

Valid

Tongtherm, Nabhitabhata & Tuanapaya

Triassic

Chaiburi Formation

 Thailand

A member of the family Ceratitidae.

Evopharciceras[20]

Gen. et sp. et comb. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany
 Morocco

A member of Agoniatitida belonging to the family Pharciceratidae. The type species is E. formosum; genus also includes "Pharciceras" evolvens Bockwinkel, Becker & Ebbighausen (2009).

Extropharciceras metallicum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Galfettites orientalis[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Galfettitidae.

Glyptophiceras mirum[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Xenoceltitidae.

Gonioclymenia junggarensis[21]

Sp. nov

In press

Zong, Becker & Ma

Devonian (Famennian)

Hongguleleng Formation

 China

Hadrothisbites hanwangensis[22]

Sp. nov

In press

Mietto et al.

Late Triassic (Carnian)

Ma'antang Formation

 China

Holcodiscus pseudocaillaudi[16]

Sp. nov

Valid

Vermeulen et al.

Early Cretaceous (Barremian)

 France

A member of the family Holcodiscidae.

Hulenites kitamurai[23]

Sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Inaptychoceras[23]

Gen. et sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Genus includes new species I. koikei.

Kislovodskoceras[24]

Gen. et sp. nov

Valid

Polkovoy

Early Cretaceous (Aptian)

 Russia

A member of the family Desmoceratidae. The type species is K. kislovodskense.

Kurnubiella fallax[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Upper Dhruma Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Laevisimoceras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is L. inornatum.

Linguatornoceras asbeckense[18]

Sp. nov

Valid

Korn

Devonian (Frasnian)

 Germany

A member of the family Tornoceratidae.

Lunupharciceras kochi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Lutites[1]

Gen. et 5 sp. et comb. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae. The type species is L. lyriformis; genus also includes new species L. alius, L. paucis, L. plicatus and L. profundus, as well as Alibashites stepanovi Korn & Ghaderi in Korn et al. (2015).

Lytogyroceras tuberculatum[11]

Sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Lytogyroceratinae.

Maenioceras ornatum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Maenioceratidae.

Nejdia duartei[25]

Sp. nov

Valid

Goy & Comas-Rengifo

Early Jurassic (Toarcian)

A member of the family Hildoceratidae.

Obtusicostites gigas[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Upper Dhruma Formation

 Saudi Arabia

A member of the family Perisphinctidae.

Ophimullericeras[26]

Gen. et sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Ceratitida belonging to the group Meekocerataceae and the family Mullericeratidae. The type species is O. paullae.

Pachyerymnoceras arabicum[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Tuwaiq Mountain Limestone Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Pachyerymnoceras magnum[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Callovian)

Tuwaiq Mountain Limestone Formation

 Saudi Arabia

A member of the family Pachyceratidae.

Palaeophyllites mishkini[7]

Sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Palaeophyllitidae.

Paracadoceras vasily[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Paracochloceras nunminensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Parahoplites inaensis[23]

Sp. nov

In press

Matsukawa

Early Cretaceous (Aptian)

Todai Formation

 Japan

Parasimoceras[11]

Gen. et 2 sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. venturii; genus also includes P. pseudostenonis.

Paratirolites aduncus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites baghukensis[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites lanceolobatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites robustus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Paratirolites rubens[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

Peripleurites gabbensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

A member of Choristocerataceae belonging to the family Rhabdoceratidae.

Pharciceras beyrichi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pharciceras ferrum[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pharciceras kruegeri[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Placites heggi[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

A member of the family Gymnitidae.

Planisimoceras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Parasimoceratinae. The type species is P. costotuberculatum.

Pluripharciceras ahlburgi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Pronjaites yolkinensis[28]

Sp. nov

Valid

Mitta

Early Cretaceous

 Russia

Pseudogalfettites[7]

Gen. et sp. nov

Valid

Smyshlyaeva & Zakharov

Early Triassic (Olenekian)

 Russia

A member of the family Galfettitidae. Genus includes new species P. sakhnoi.

Psiloceras pseudocostosum[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Retrotornoceras juxi[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Rhacophyllites mulleri[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Rhacophyllites volcanoensis[5]

Sp. nov

Valid

Taylor, Guex & Lucas

Late Triassic (Rhaetian)

Gabbs Formation

 United States
( Nevada)

Rollierites biplicatum[29]

Sp. nov

Valid

Jain & Mazur

Middle Jurassic (Callovian)

 Poland

A member of the family Pachyceratidae.

Sandbergeroceras archiaci[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Triainoceratidae.

Saudicostites[3]

Gen. et sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bathonian)

Dhruma Formation

 Saudi Arabia

A member of Perisphinctoidea of uncertain phylogenetic placement. The type species is S. arabicus.

Shevyrevites corrugatus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Xenodiscidae.

Sigaloceras fundator[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Sigaloceras kiselevi[27]

Sp. nov

Valid

Gulyaev in Gulyaev & Ippolitov

Middle Jurassic (Callovian)

 Ukraine

Simocosticeras[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. biornatum.

Sinotropites[22]

Gen. et sp. nov

In press

Mietto et al.

Late Triassic (Carnian)

Ma'antang Formation

 China

Genus includes new species S. sichuanensis.

Sonninia (Euhoploceras) crassicostae[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Lower Dhruma Formation

 Saudi Arabia

A member of the family Sonniniidae.

Sonninia (Euhoploceras) turbator[3]

Sp. nov

Valid

Énay & Mangold

Middle Jurassic (Bajocian)

Lower Dhruma Formation

 Saudi Arabia

A member of the family Sonniniidae.

Sonoraceras[30]

Gen. et sp. nov

In press

Samaniego-Pesqueira, Moreno-Bedmar & Álvarez-Sánchez

Early Cretaceous (Aptian)

 Mexico

Genus includes new species S. tepachensis.

Stenopharciceras lotzi[20]

Sp. nov

Valid

Korn & Bockwinkel

Devonian (Givetian)

Roteisenstein Formation

 Germany

A member of Agoniatitida belonging to the family Pharciceratidae.

Stoliczkaia (Lamnayella) duboisi[31]

Sp. nov

Valid

Kennedy & Latil

Late Cretaceous (Cenomanian)

 France

A member of the family Lyelliceratidae.

Stoyanowites parallelus[1]

Sp. nov

Valid

Korn & Hairapetian in Korn et al.

Permian (Changhsingian)

Hambast Formation

 Iran

A member of the family Dzhulfitidae.

Strenoceras djangurense[32]

Sp. nov

Valid

Mitta

Middle Jurassic (Bajocian)

 Russia

Sulcicostites[11]

Gen. et sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Simoceratinae. The type species is S. costatus.

Timorites dendroides[33]

Sp. nov

Valid

Korn & Ghaderi in Tabrizi et al.

Permian (Wuchiapingian)

Julfa Formation

 Iran

A member of the family Cyclolobidae.

Tornoceras aequilobum[8]

Sp. nov

Valid

Söte et al.

Devonian (Frasnian)

Sand Formation

 Germany

A member of the family Tornoceratidae.

Tornoceras fundibulum[18]

Sp. nov

Valid

Korn

Devonian (Famennian)

A member of the family Tornoceratidae.

Tornoceras occasus[34]

Sp. nov

Valid

Korn

Late Devonian

 Morocco

A member of the family Tornoceratidae.

Tornoceras ortus[34]

Sp. nov

Valid

Korn

Late Devonian

 Morocco

A member of the family Tornoceratidae.

Tornoceras vretterense[18]

Sp. nov

Valid

Korn

Devonian (Givetian)

A member of the family Tornoceratidae.

Tropaeum evolutum[35]

Sp. nov

In press

Lehmann & Bayliss

Early Cretaceous (Aptian)

Hythe Formation

 United Kingdom

Tropidoceras komarovi[36]

Sp. nov

Valid

Zaitsev

Early Jurassic

Crimean Peninsula

Vicininodiceras dalriatense[12]

Sp. nov

Valid

Simms & Edmunds

Early Jurassic

 United Kingdom

Volanoceras cuneospinatum[11]

Sp. nov

Valid

Nannarone & Bilotta

Late Jurassic (Tithonian)

 Italy

A member of Perisphinctoidea belonging to the family Simoceratidae and the subfamily Volanoceratinae.

Waehneroceras morawetzi[10]

Sp. nov

Valid

Kment

Early Jurassic (Hettangian)

Wordieoceras mullenae[26]

Sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Ceratitida belonging to the group Meekocerataceae and the family Ophiceratidae.

Research[edit]

  • A modern review of the palaeobiology of heteromorph ammonoids is published by Hoffmann et al. (2021), including details of their anatomy, buoyancy, locomotion, predators, diet, palaeoecology, and extinction.[37]
  • A study on the impact of the Permian–Triassic extinction event on the morphological diversity of ammonites is published by Dai, Korn & Song (2021).[38]
  • A study on the morphology and ontogeny of whorl profiles of the late Anisian ceratitid ammonites is published by Bischof et al. (2021).[39]
  • An exceptionally preserved specimen of Sigaloceras enodatum, providing information on the anatomy of ammonite muscles, is described from the Middle Jurassic (Callovian) Kellaways Sand Member of Colne Gravel (Gloucestershire, United Kingdom) by Cherns et al. (2021), who evaluate the implications of this specimen for the knowledge of the swimming mechanism of ammonites.[40]
  • Soft parts of a perisphinctid belonging to the genus Subplanites, separated from the conch either taphonomically or during a failed predation, are described from the Tithonian conservation deposits of Wintershof (southern Germany) by Klug et al. (2021).[41]
  • The first physical compression experiments on model ammonite septa are performed by Johnson et al. (2021), who attempt to determine whether the increasingly complex fractal margins of ammonite septa might have evolved to increase resistance to shell-crushing predators.[42]
  • A study aiming to determine the function of complex septa in the ammonite shell, providing evidence of a relationship between septal complexity and capacity for liquid retention within chambers of the shell, likely improving buoyancy regulation, is published by Peterman et al. (2021).[43]
  • Two specimens of Peltoceras athleta having both female and male features are described from Callovian limestone beds in Méron near Montreuil-Bellay (France) by Frau & Boursicot (2021), who interpret these specimens as cases of intersexuality in ammonites.[44]
  • Soft tissue remains preserved within the conch of a specimen of Proleymeriella schrammeni from the Albian of Germany are described by Hoffmann et al. (2021).[45]
  • A study on the arrangement in space and probable nature of hook-like structures found with specimens of Rhaeboceras halli is published by Smith et al. (2021), who interpret these structures as representing part of the brachial crown armature, and propose a hypothetical reconstruction of an ammonite brachial crown.[46]
  • A study on the systematics and phylogeny of the genus Aegocrioceras is published by Weinkauf, Hoffmann & Wiedenroth (2021).[47]
  • A study on the hydrostatic properties of Baculites compressus, and on their implications for the knowledge of the swimming capabilities and ecology of this ammonite and orthoconic cephalopods in general, is published by Peterman & Ritterbush (2021).[48]
  • Revision of the fossil material of Parapuzosia seppenradensis, including historic specimens from Europe and the United States and new material from England and Mexico, and a study on the ontogeny, distribution and evolution of this ammonite, is published by Ifrim et al. (2021).[49]
  • A study on the habitat depth preferences of Hoploscaphites constrictus from the European Boreal Chalk Sea, as indicated by stable isotope thermometry of aptychi and by data from predation marks on the scaphitid moulds, is published by Machalski et al. (2021).[50]
  • A study on the relationship between sutural complexity and body size in Cretaceous ammonites is published by Pérez-Claros (2021).[51]

Other cephalopods[edit]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acrocoelites ippolitae[52]

Sp. nov

Valid

Weis in Weis et al.

Middle Jurassic (Aalenian)

 Germany

A belemnite.

Anglonautilus hewaidyi[53]

Sp. nov

In press

Moneer, Bazeen & Kassab

Late Cretaceous (Maastrichtian)

 Egypt

A nautiloid cephalopod.

Angulithes monqarensis[53]

Sp. nov

In press

Moneer, Bazeen & Kassab

Late Cretaceous (Maastrichtian)

 Egypt

A nautiloid cephalopod.

Arionoceras tokushimaense[54]

Sp. nov

Valid

Niko

Silurian (Ludlow)

Suberidani Group

 Japan

Bactroceras fluvii[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Dapingian)

Valhallfonna Formation

 Norway

Boreopeltis ifrimae[56]

Sp. nov

In press

Fuchs & Stinnesbeck

Late Cretaceous (Turonian)

 Mexico

Buttsoceras buldrebreenense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Troedssonellidae.

Cycloplectoceras hinlopense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Tarphyceratidae.

Cyclostomiceras profilstrandense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Cyclostomiceratidae.

Deltoceras beluga[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Cyclostomiceratidae.

Eosomichelinoceras borealis[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Baltoceratidae.

Eosomichelinoceras submedius[57]

Sp. nov

Valid

Evans et al.

Ordovician (Darriwilian)

Pelmis Formation

 Iran

A member of the family Baltoceratidae.

Ethanoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is E. solitudines.

Gzheloceras aisenvergi[58]

Sp. nov

Valid

Dernov

Carboniferous (Serpukhovian)

Samara Formation

 Ukraine

A member of the family Tainoceratidae.

Hemichoanella occulta[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Hinlopoceras[55]

Gen. et 2 sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is H. tempestatis; genus also includes H. venti.

Jovellania praetermissa[59]

Sp. nov

Valid

Brauckmann & Koch

Devonian (Givetian)

Massenkalk Formation

 Germany

Knightoceras extorris[58]

Sp. nov

Valid

Dernov

Carboniferous (Pennsylvanian)

 Ukraine

A member of the family Koninckioceratidae.

Lawrenceoceras ebenus[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Lawrenceoceras larus[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Litoceras profilbekkenense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Mojsvaroceras gianii[60]

Sp. nov

Valid

Pieroni & Prinoth

Middle Triassic (Anisian)

 Italy

A nautiloid.

Nyfrieslandoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is N. bassleroceroides.

Olenidslettoceras[55]

Gen. et sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of Oncocerida belonging to the family Phthanoncoceratidae. The type species is O. farmi.

Paracenoceras waageni[61]

Sp. nov

Valid

Halder

Middle Jurassic (Callovian)

Chari Formation

 India

A member of the family Paracenoceratidae.

Planetoceras yefimenkoi[58]

Sp. nov

Valid

Dernov

Carboniferous (Bashkirian)

Mospinka Formation

 Ukraine

A member of the family Koninckioceratidae.

Proterocameroceras valhallfonnense[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

Protocycloceras minor[55]

Sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A member of the family Troedssonellidae.

Rhynchorthoceras yizanense[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Sinoceras complexum[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Sorosoceras[57]

Gen. et sp. nov

Valid

Evans et al.

Ordovician (Darriwilian)

Pelmis Formation

 Iran

A member of the family Geisonoceratidae. The type species is S. castellum.

Svalbardoceras[55]

Gen. et 2 sp. nov

Valid

Kröger & Pohle

Ordovician (Floian)

Valhallfonna Formation

 Norway

A relative of Rioceras. The type species is S. sterna; genus also includes S. skua.

Tyrioceras longicameratum[62]

Sp. nov

Valid

Fang in Fang et al.

Ordovician

 China

A member of Lituitida.

Xiaohenautilus mulleni[26]

Sp. nov

Valid

Jenks et al.

Early Triassic

Dinwoody Formation

 United States
( Nevada)

A member of Nautilida belonging to the group Trigonoceratoidea and the family Grypoceratidae.

Research[edit]

  • New fossil material with shell features that characterize early cephalopods is described from the Cambrian (Terreneuvian) Bonavista Formation (Canada) by Hildenbrand et al. (2021), who interpret this finding as possibly representing the earliest cephalopods known to date, potentially pushing the origin of cephalopods back in time by about 30 million years, to the time before the first occurrence of euarthropods.[63]
  • Two specimens of Syrionautilus libanoticus preserving soft parts, including a specimen showing an imprint of the conch, are described from the Cenomanian of Hadjoula (Lebanon) by Klug et al. (2021).[64]
  • A study on the diversity patterns and spatial structure of belemnite assemblages in Western Tethys Ocean during the Early Jurassic is published by Neige, Weis & Fara (2021).[65]
  • A study on morphological changes of belemnites across the Pliensbachian-Toarcian boundary event, based on data from the Peniche GSSP section (Portugal), is published by Nätscher et al. (2021).[66]
  • A study on chemistry, organization and genesis of circular structures (superficially resembling chromatophores) preserved in coleoid cephalopod specimens from the Jurassic of Germany and the Cretaceous of Lebanon is published by Klug et al. (2021).[67]
  • A well-preserved soft-body imprint of a fossil squid is described from the lower Oligocene of the Krasnodar region (Russia) by Mironenko et al. (2021), who interpret this finding as the first unquestionable representative of Teuthida in the fossil record known to date, and report preservation of fish remains in the stomach contents of this squid, indicative of its piscivorous diet.[68]
  • A specimen of Jeletzkyteuthis coriacea holding a specimen of Parabelopeltis flexuosa in its arms, with the apex very close to its jaws, is described from the Toarcian Posidonia Shale (Germany) by Klug et al. (2021), who interpret this finding as a case of frozen predatory behaviour of two different octobrachian species.[69]
  • Redescription of the gladius of the holotype specimen of Necroteuthis hungarica is published by Košťák et al. (2021), who reinterpret this species as a member of the family Vampyroteuthidae linking Mesozoic loligosepiids with extant Vampyroteuthis.[70]
  • Klompmaker & Landman (2021) describe the oldest drill holes produced by octopodoids in lucinid bivalve specimens from the Campanian of South Dakota, indicating that the drilling habit evolved early in the evolutionary history of Octopodoidea.[71]

Gastropods[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acirsa ambrosii[72]

Sp. nov

Valid

Brunetti & Cresti

Pliocene

 Italy

A species of Acirsa.

Acirsa (Notacirsa) texana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Acirsa.

Acteon pseudotypica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Acteon.

Acutitomaria[74]

Gen. et sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida. Genus includes new species A. kustatscherae.

Alatostrombus[75]

Gen. et sp. et comb. nov

Valid

Raven

Miocene

 Indonesia
 Malaysia

A member of the family Strombidae. Genus includes new species A. acutalatus, as well as "Strombus" sedanensis Martin (1899).

Alvania absuturalis[76]

Sp. nov

Valid

Chirli & Forli

Pliocene

 Italy

A species of Alvania.

Alvania ameliae[76]

Sp. nov

Valid

Chirli & Forli

Pleistocene

 Italy

A species of Alvania.

Alvania benestarensis[76]

Sp. nov

Valid

Chirli & Forli

Miocene

 Italy

A species of Alvania.

Alvania cerreti[77]

Sp. nov

Valid

Gardella et al.

Pliocene

 Italy

A species of Alvania.

Alvania pluricosticillata[76]

Sp. nov

Disputed

Chirli & Forli

Plio-Pleistocene

 Italy

Originally described as a species of Alvania. Argued to be a junior synonym of the extant taxon Crisilla gagliniae by Amati & Oliverio (2024).[78]

Amaea (Confusiscala) paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Ammonicera danieli[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A species of Ammonicera.

Ammonicera rolani[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A species of Ammonicera.

Ampezzogyra angulata[80]

Sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Stuoraxidae.

Amphidromus ubaldii[81]

Sp. nov

Valid

Dharma

Late Pliocene

 Indonesia

A species of Amphidromus.

Amplitomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" spuria Münster (1841).

Anacithara akisi[82]

Sp. nov

Valid

Kolokotronis

Pliocene

 Cyprus

A species of Anacithara.

Anceps siminescui[83]

Nom. nov

Valid

Harzhauser

Middle-late Miocene

 Moldova

A Trochidae member;
a replacement name for Trochus semistriatus Siminescu & Barbu (1940).

Angulatella[80]

Gen. et sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Prostyliferidae. The type species is A. bizzarinii.

Antithala[84]

Gen. et comb. et 2 sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Bosnia and Herzegovina
 Hungary
 Poland
 Romania
 Ukraine

A member of the family Costellariidae. The type species is "Mitra" sturi Hoernes & Auinger (1880); genus also includes "Mitra" neugeboreni Hoernes & Auinger (1880), as well as new species A. claviformis and A. filipescui.

Apiocypraea cerretensis[85]

Sp. nov

Valid

Fehse et al.

Pliocene (Zanclean)

 Italy

A member of the family Eocypraeidae.

Aplus abessensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus grandiculus[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus littoralis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aplus pseudoaquitanensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Aplus.

Aquilofusus? antiqua[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Ascheria canni[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae.

Asiaharpa[88]

Gen. et sp. nov

Valid

Raven

Miocene

 Malaysia

A member of the family Harpidae. Genus includes new species A. sarawakiana.

Attenuata? danica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Attenuata.

Auingeria[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Germany
 Hungary

A member of the family Columbellidae. The type species is "Columbella" bellardii Hörnes (1852); genus also includes "Columbella (Anachis)" austriaca Hoernes & Auinger (1880) and "Pyrene (Anachis) bellardii" grussbachensis Csepreghy-Meznerics (1969; raised to the rank of a separate species Auingeria grussbachensis).

Axonella[90]

Gen. et comb. et sp. nov

Valid

Pacaud, Lhomme & Renaud

Eocene (Ypresian)

 France
 Spain

A member of the family Hemisinidae. The type species is "Cerithium" palense Rouault (1850); genus also includes "Potamides" granosus Doncieux (1908), as well as new species A. falsiacum.

Babylonia chattica[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Babylonia.

Babylonia edigittenbergeri[92]

Sp. nov

Valid

Raven

Miocene

Miri Formation

 Malaysia

A species of Babylonia.

Babylonia sarawakiana[92]

Sp. nov

Valid

Raven

Miocene

Miri Formation

 Malaysia

A species of Babylonia.

Bandellina compacta[80]

Sp. nov

Valid

Nützel & Hausmann in Hausmann et al.

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A member of the family Cornirostridae.

Barycypraea abbasi[93]

Sp. nov

Valid

Dovesi & Parsons

Miocene (Serravallian)

Nyalindung Formation

 Indonesia

A species of Barycypraea.

Barycypraea alessandrodovesi[93]

Sp. nov

Valid

Dovesi & Parsons

Miocene (Serravallian)

Nyalindung Formation

 Indonesia

A species of Barycypraea.

Bela pediensis[94]

Sp. nov

Valid

Kolokotronis

Pliocene

 Cyprus

A species of Bela.

Bellacolumbella[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Hungary
 Romania

A member of the family Columbellidae. The type species is "Columbella (Nitidella)" karreri Hoernes & Auinger (1880); genus also includes "Columbella (Nitidella)" embryonalis Boettger (1902) and "Columbella (Nitidella)" katharinae Hoernes & Auinger (1880).

Bellardithala[84]

Nom. et 4 sp. nov

Valid

Harzhauser & Landau

Miocene and Pliocene

Baden Formation
Dej Formation

 Austria
 Czech Republic
 Italy
 Poland
 Romania
 Ukraine
 Hungary?

A member of the family Costellariidae; a replacement name for Micromitra Bellardi (1888). The type species is "Voluta" obsoleta Brocchi (1814); genus also includes new species B. kovaci, B. baluki, B. fedosovi and B. dacica.

Bieleraxis[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species B. discoidea and B. altis.

Bithynia disregularis[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia obliquus[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia parainclinimarga[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bithynia paramonolithic[95]

Sp. nov

In press

Li et al.

Oligocene

Yehucheng Formation

 China

A species of Bithynia.

Bittium prekoeneni[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Bittium.

Bulimulus frenguellii[96]

Sp. nov

Valid

Cabrera et al.

Late Cretaceous

Queguay Formation

 Uruguay

A species of Bulimulus.

Burminella[97]

Gen. et sp. nov

In press

Balashov, Perkovsky & Vasilenko

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A land snail possessing morphological traits intermediate between members of the family Pupinidae and other members of Cyclophoroidea. Genus includes new species B. artiukhini.

Bursa transeuntis[98]

Sp. nov

Valid

Forli & Smriglio

Miocene and Pliocene

 Hungary
 Italy
 Poland

A species of Bursa.

Caecum texanum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Caecum.

Califrapana[99]

Gen. et comb. nov

Valid

Powell & Houart

Late Oligocene to early Miocene

Vaqueros Formation

 Mexico
 United States
( California)

A member of the family Muricidae. The type species is "Purpura" vaquerosensis Arnold (1907).

Calyptraphorus velusus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Camponaxis bandeli[100]

Sp. nov

Valid

Pieroni, Monari & Todd

Late Triassic (Carnian)

San Cassiano Formation

 Italy

A Tofanellidae member

Campylacrum texanum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Canariella (Majorata) gerti[101]

Sp. nov

Valid

Groh

Quaternary

 Canary Islands

A species of Canariella.

Cancellotomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" subcancellata d'Orbigny (1850).

Cancilla nanostriatula[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bulgaria
 Hungary

A species of Cancilla.

Cancilla wagreichi[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Cancilla.

Capulus gelus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Capulus.

Cassis kalmani[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A species of Cassis.

Cassis rasae[104]

Sp. nov

Valid

Petuch & Berschauer

Pleistocene

 United States
( Florida)

A species of Cassis.

Cassis viliusi[104]

Sp. nov

Valid

Petuch & Berschauer

Pleistocene

 United States
( Florida)

A species of Cassis.

Cenorhytis[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species C. groesbecki.

Ceratopea? moridunensis[105]

Sp. nov

Valid

Ebbestad & Cope

Ordovician (Floian)

 United Kingdom

A member of the family Raphistomatidae.

Cerithiella acuta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cerithiella.

Cerithiscala parvibrazosa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cerithiscala tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Chemnitzia silvai[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Chemnitzia.

Chevallieria texanopsis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Chicomurex parvus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicomurex.

Chicoreus javanus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Chicoreus martini[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Chicoreus solidus[107]

Sp. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Chicoreus.

Clava (Semivertagus) scriba[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Clavatula hirmetzli[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Clavatula santhai[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Clavatula szekelyhidiae[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Clavatula.

Cochlespira (Tahusyrinx) laevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cochlespira.

Conilithes herodus[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece

A species of Conilithes.

Conradconfusus nodulinus[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Conradconfusus texanus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Conus (Kalloconus) asterousiaensis[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece
 Italy

A species of Conus.

Conus (Kalloconus) helladicus[109]

Sp. nov

Valid

Psarras, Koskeridou & Merle

Miocene (Tortonian)

 Greece

A species of Conus.

Cordieria biouesensis[111]

Sp. nov

Valid

Pacaud

Paleocene (Thanetian)

 France

A species of Cordieria.

Coronia argilla[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Costoanachis venzoi[89]

Nom. nov

Valid

Harzhauser & Landau

Miocene

 Italy

A species of Costoanachis; a replacement name for Anachis (Costoanachis) subcorrugata Venzo & Pelosio (1963).

Crassaseila[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species C. ripleyana and C. paleocenica.

Crassispira plioibericostricta[112]

Sp. nov

Valid

Brunetti & Forli

Pliocene (Zanclean)

 Spain

A species of Crassispira.

Crassispira pseudoglabra[112]

Sp. nov

Valid

Brunetti & Forli

Pliocene (Piacenzian)

 Italy

A species of Crassispira.

Crenaturricula minuta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Creonella hebetertia[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cretatortulosa cretakachinensis[113]

Sp. nov

In press

Yu, Salvador & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Cretatortulosa gignens[114][115][116]

Sp. nov

In press

Jochum, Yu & Neubauer

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Pupinidae.

Cryptochorda paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cyathopoma pingyiensis[117]

Sp. nov

In press

Yu et al.

CretaceousPaleogene transition

Pingyi Basin

 China

A species of Cyathopoma.

Cylichna (Cylichnopsis) elliptica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cylichna.

Cymatiella dulaii[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A species of Cymatiella.

Cymatiella tzankovi[103]

Nom. nov

Valid

Vicián & Kovács

Paleocene

 Bulgaria

A species of Cymatiella; a replacement name for Eutritonium (Sassia) rutoti Tzankov (1940).

Cymatium danianum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cymatium.

Cyphoma guraboensis[118]

Sp. nov

Valid

Groves & Landau

Early Pliocene

Gurabo Formation

 Dominican Republic

A species of Cyphoma.

Cypraeorbis alisonkayae[118]

Sp. nov

Valid

Groves & Landau

Miocene

 Dominican Republic

A species of Cypraeorbis.

Cyprioconcha[87]

Gen. et sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae. The type species is C. robertsoni.

Defensina[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Bosnia and Herzegovina
 Bulgaria
 Romania

A member of the family Columbellidae. The type species is "Columbella (Mitrella)" bucciniformis Hoernes & Auinger (1880).

Dermomurex wanneri[107]

Nom. nov

Valid

Merle, Landau & Breitenberger

Miocene

 Indonesia

A species of Dermomurex; a replacement name for Murex acuticostatus Wanner & Hahn (1935).

Desbruyeresia kambiaensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Desbruyeresia kinousaensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Desbruyeresia memiensis[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A species of Desbruyeresia.

Discotectonica paracutissima[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Discotectonica.

Doliella parnitens[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Doliella.

Domiporta amoena[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Austria
 Bulgaria
 Romania

A species of Domiporta.

Domiporta pulchra[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Domiporta.

Domiporta turpis[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria

A species of Domiporta.

Ederazyga[100]

Gen. et sp. et comb. nov

Valid

Pieroni, Monari & Todd

Late Triassic (Carnian to Rhaetian)

Nayband Formation
San Cassiano Formation
Zu Limestone

 Iran
 Italy
 Slovenia

A possible Zygopleuridae member.
The type species is E. fanchini;
genus also includes "Cerithium"? lateplicatum Klipstein (1843).

Ellobium mizutanii[119]

Sp. nov

Valid

Kawase & Ichihara

Early Miocene

Akeyo Formation

 Japan

A species of Ellobium.

Entemnotrochus kathiawarensis[120]

Sp. nov

Valid

Bose, Das & Mondal

Miocene (Burdigalian)

 India

A species of Entemnotrochus.

Eoacteon paracutissima[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eoancilla lapicidina[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eocithara (?) laurencei[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eopleurotoma atacica[111]

Nom. nov

Valid

Pacaud

Eocene (Ypresian)

 France

A species of Eopleurotoma; a replacement name for Pleurotoma (Eopleurotoma) romani Doncieux (1908).

Eotrivia pacaudi[121]

Sp. nov

Valid

Fehse

Eocene

 Italy

Eoturris fallsensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Eoworthenia frydai[74]

Nom. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida; a replacement name for Worthenia rarissima Barrande.

Episcomitra antibellardii[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Episcomitra.

Episcomitra leopoldiana[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Czech Republic
 Poland

A species of Episcomitra.

Episcomitra missile[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Grund Formation

 Austria
 Hungary

A species of Episcomitra.

Episcomitra neubaueri[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Episcomitra.

Episcomitra pontonxensis[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Episcomitra.

Episcomitra pseudoincognita[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania
 Ukraine

A species of Episcomitra.

Episcomitra vermeuleni[122]

Sp. nov

Valid

Pacaud

Eocene (Priabonian)

 Ukraine

A species of Episcomitra.

Eulimella ariejansseni[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Eulimella.

Euthema myanmarica[123]

Sp. nov

Valid

Balashov

Cretaceous

Hkamti amber

 Myanmar

A species of Euthema.

Euthria mellianensis[86]

Sp. nov

Valid

Lozouet

Miocene

 France

A species of Euthria.

Euthria tarusatensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Euthria.

Euthriofusus argillensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

Falsifusus problematicus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Fedosovia[84]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Czech Republic
 Italy
 Romania

A member of the family Costellariidae. The type species is "Mitra (Callithea)" fuchsi Hoernes & Auinger (1880).

Fraudiziba[102]

Gen. et nom. et sp. nov

Valid

Harzhauser & Landau

Miocene and Pliocene

Karaman Basin

 Austria
 Bosnia and Herzegovina
 Czech Republic
 Hungary
 Italy
 Poland
 Romania
 Slovakia
 Turkey
 Ukraine

A member of the family Mitridae. The type species is F. paratethyca (a replacement name for Mitra austriaca Csepreghy-Meznerics, 1950); genus also includes new species F. ottomanica.

Fulgurofusus spinosus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Fulgurofusus.

Fusulculus texanus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Fusulculus.

Galba prima[124]

Sp. nov

Valid

Yu, Neubauer & Jochum

Cretaceous

Burmese amber

 Myanmar

A species of Galba.

Gemmula bearrizensis[111]

Nom. nov

Valid

Pacaud

Eocene (Priabonian)

 France

A species of Gemmula; a replacement name for Drillia pellati Boussac (1911).

Gemmula osca[111]

Sp. nov

Valid

Pacaud

Eocene (Bartonian)

 Spain

A species of Gemmula.

Gentilifulgur[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species G. praemortoni.

Gibbula lovellreevei[83]

Nom. nov

Valid

Harzhauser

Pliocene

 United Kingdom

A Trochidae member;
a replacement name for Trochus (Gibbula) reevei Harmer (1923).

Gibbula pizzolatoi[125]

Sp. nov

Valid

Spadini

Pliocene

 Italy

A species of Gibbula.

Gibbula steiningeri[83]

Nom. nov

Valid

Harzhauser

Early Miocene

 Austria

A Trochidae member;
a replacement name for Trochus amedei bicincta Schaffer (1912).

Gibbula tavanii[83]

Nom. nov

Valid

Harzhauser

Miocene

 Libya

A Trochidae member;
a replacement name for Gibbula minima Tavani (1939).

Gibbuliculus[83]

Gen. et comb. et nom. nov

Valid

Harzhauser

Oligocene to Pleistocene

 Europe

A Trochidae member;
The type species is "Gibbula" brebioni Landau, Van Dingenen & Ceulemans (2017).
Genus includes G. saccoi (a replacement name for Gibbula protumida Sacco, 1896).

"Gibbulinella" sandbergeri[126]

Nom. nov

Valid

Harzhauser & Neubauer

Eocene

 Italy

A member of the family Streptaxidae; a replacement name for Pupa simplex Sandberger (1870).

Haedropleura dellabellai[72]

Sp. nov

Valid

Brunetti & Cresti

Pliocene

 Italy

A species of Haedropleura.

Hammatospira cancellata[127]

Sp. nov

Valid

Mazaev

Early Carboniferous

 Kazakhstan

Hespererato canae[118]

Sp. nov

Valid

Groves & Landau

Late Miocene and early Pliocene

Cercado Formation

 Dominican Republic

A species of Hespererato.

Hespererato praeclusa[118]

Sp. nov

Valid

Groves & Landau

Early–middle Miocene

Baitoa Formation

 Dominican Republic

A species of Hespererato.

Hexaplex nascium[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Hexaplex.

Hirsuticyclus canaliculatus[128]

Sp. nov

In press

Yu

Cretaceous

Burmese amber

 Myanmar

A member of the family Cyclophoridae.

Hochheimia[126]

Nom. nov

Valid

Harzhauser & Neubauer

Oligocene

 Germany

A member of the family Hygromiidae; a replacement name for Palaeotrichia Nordsieck (2014).

Hokkaidoconcha morisseaui[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of the family Hokkaidoconchidae.

Holospira? fallsensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Holospira.

?Hydrocena praecursor[129]

Sp. nov

In press

Yu & Neubauer

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Hydrocenidae.

Jponsia lacetania[130]

Sp. nov

Valid

Pacaud & Goret

Eocene (Ypresian)

 Spain

A member of the family Pachychilidae.

Kapalmerella gelus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Keepingia elongata[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

Laevispira[75]

Gen. et comb. et sp. nov

Valid

Raven

Miocene and Pliocene

 Brunei
 India
 Indonesia

A member of the family Strombidae. Genus includes "Dolomena" bruneiensis Harzhauser, Raven & Landau (2018), "Strombus" glaber Martin (1879), "Strombus" karikalensis Cossmann (1903), "Strombus" triangulatus Martin (1879) and "Strombus" varinginensis Martin (1899), as well as new species L. seriaensis.

Laevitrivia[131]

Gen. et comb. et 5 sp. nov

Valid

Fehse & Vicián

Miocene and Pliocene

 France
 Italy

A member of the family Triviidae. The type species is "Cypraea" pisolina Lamarck, (1810); genus also includes new species L. annae, L. bertaccinii, L. pliopisolina, L. pseudocalva and L. ventricostata.

Latirus? asperpellis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Latirus.

Laubella subsulcata[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Lineacingulum[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" texturata Münster (1841).

Lineaetomaria[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" decorata Münster (1841).

Liostomia wilvanderstoelae[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Liostomia.

Littorina elongata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Littorina.

Littorina (Prosthenodon) incertus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Littorina.

Lupira danianum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Lutema gracilis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Lyria elongata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lyria.

Lyria tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lyria.

Malea hyaducki[104]

Sp. nov

Valid

Petuch & Berschauer

Pliocene

 United States
( Florida)

A species of Malea.

Martaia[89]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Austria
 Bulgaria
 Czech Republic
 Hungary
 Romania
 Ukraine

A member of the family Columbellidae. The type species is "Columbella (Anachis)" dujardini Hoernes & Auinger (1880); genus also includes "Columbella (Anachis)" zitteli Hoernes & Auinger (1880).

Mathilda (Echinimathilda) duogenta[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Mathilda.

Melongena pontonxensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Melongena.

Metula bulbiformis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula nana[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula pseudomajor[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Metula rivulisensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Metula.

Mimospira llangynogensis[105]

Sp. nov

Valid

Ebbestad & Cope

Ordovician (Floian)

 United Kingdom

A member of the family Clisospiridae.

Mitrella dacica[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Hungary
 Romania
 Slovakia

A species of Mitrella.

Mitrella demaintenonae[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation
Dej Formation

 Austria
 Romania

A species of Mitrella.

Mitrella elongatissima[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bosnia and Herzegovina

A species of Mitrella.

Mitrella fortis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Mitrella.

Mitrella tarbelliana[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Mitrella.

Mitrella viennensis[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bulgaria
 Czech Republic
 Kazakhstan
 Poland
 Romania

A species of Mitrella.

Moniquia astibiai[130]

Sp. nov

Valid

Pacaud & Goret

Eocene (Priabonian)

 Spain

A member of the family Pachychilidae.

Monoplex s.l. szakonyii[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A member of the family Cymatiidae.

Mulderia mulderi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A member of the family Pyramidellidae.

Multitrochia[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species M. tehuacana.

Nannamoria alquezae[132]

Sp. nov

Valid

Hawke

Miocene

 Australia

A species of Nannamoria.

Nannamoria cadella[132]

Sp. nov

Valid

Hawke

Miocene

Cadel Formation

 Australia

A species of Nannamoria.

Nannamoria costatum[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria flindersi[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria gnotuki[132]

Sp. nov

Valid

Hawke

 Australia

A species of Nannamoria.

Nannamoria hiscocki[132]

Sp. nov

Valid

Hawke

Pliocene

Cameron Inlet Formation

 Australia

A species of Nannamoria.

Nannamoria malonei[132]

Sp. nov

Valid

Hawke

 Australia

A species of Nannamoria.

Nannamoria persimilis[132]

Sp. nov

Valid

Hawke

Miocene

 Australia

A species of Nannamoria.

Naria praehelvola[133]

Sp. nov

Valid

Fehse & Vicián

Miocene

 Indonesia

A species of Naria.

Natica (Carinacca) kincaidensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Natica.

Nebularia soliphila[102]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria

A species of Nebularia.

Nemaspira[134]

Gen. et comb. et 2 sp. nov

Valid

Mazaev

Early Carboniferous to Late Triassic

 Belgium
 Russia
 Thailand
 United Kingdom
 United States
 China?

A member of the family Gosseletinidae. Genus includes several species formerly assigned to the genera Gosseletina and Globodoma, as well as new species N. shamaevi and N. skuini.

Nitidiclavus senesi[135]

Sp. nov

Valid

Biskupič

Miocene

 Slovakia

A member of the family Drilliidae.

Niveria callosa[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Niveria.

Niveria novosperara[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Niveria.

Nodocingulum[74]

Gen. et comb. et 2 sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida. The type species is "Pleurotomaria" coronata Münster (1841); genus also includes new species N. ernstkittli and possibly also N? turris.

Nodolargena[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A member of the new family Nodolargenidae. Genus includes new species N. frosti.

Nodovoluta[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species N. coronata.

Odostomia malagensis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Odostomia.

Oligonalia[86]

Gen. et comb. nov

Valid

Lozouet

Oligocene

 France

A new genus for "Siphonalia" riparia Lozouet (1999).

Olinderia[91]

Gen. et comb. et 3 sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A member of the family Charitodoronidae. The type species is "Charitodoron" tauzini Lozouet (1991); genus also includes new species O. lestrillensis, O. poustagnacensis and O. verdunensis.

Olivella tehuacanensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Olivella.

Ondina pinguis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Ondina.

Onustus undosus[75]

Sp. nov

Valid

Raven

Miocene

 Brunei

A species of Onustus.

Opalia tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Opalia.

Orbitestella kieli[79]

Sp. nov

Valid

Chernyshev & Goedert

Late Eocene/early Oligocene

Gries Ranch Formation

 United States
( Washington)

A member of the family Orbitestellidae.

Orthosurcula ethani[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Oxyacrum gracilis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Pagonia[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species P. spiralis.

Palaeorhaphis? laevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Paleofusus[73]

Gen. et 3 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species P. bellus, P. parvicrassus and P. elegantissima.

Papuliscala? acus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Papuliscala.

Papuliscala keani[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Parasassia vargai[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A member of the family Cymatiidae.

Parthenina pacaudi[106]

Nom. nov

Valid

Landau & Micali

Miocene

 France

A species of Parthenina; a replacement name for Pyrgulina (Parthenina) dollfusi Cossmann (1921).

Parvisipho? spiralis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Paskentana dixoni[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of Abyssochrysoidea belonging to the family Paskentanidae.

Paskentana xenophontosi[87]

Sp. nov

Valid

Kaim et al.

Late Cretaceous (Cenomanian to Turonian)

 Cyprus

A member of Abyssochrysoidea belonging to the family Paskentanidae.

Perotrochus bermotiensis[120]

Sp. nov

Valid

Bose, Das & Mondal

Oligocene

 India

A species of Perotrochus.

Perrona harzhauseri[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Perrona.

Perrona nemethi[108]

Sp. nov

Valid

Kovács & Vicián

Middle Miocene

Lajta Limestone Formation

 Hungary

A species of Perrona.

Personopsis merlei[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A species of Personopsis.

Pfefferiola[126]

Nom. nov

Valid

Harzhauser & Neubauer

Eocene

 Italy

A member of the family Streptaxidae; a replacement name for Oppenheimiella Pfeffer (1930).

Phos estotiensis[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Phos.

Pleurotomella irminonvilla[111]

Nom. nov

Valid

Pacaud

Eocene (Bartonian)

 France

A species of Pleurotomella; a replacement name for Pleurotomella cossmanni Morellet & Morellet (1946).

Pliciscala obliqua[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Poirieria (Paziella) delera[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Polinices tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Polinices.

Popenoeum tabulatum[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Pressulasphaera[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" pamphilus d'Orbigny (1850).

Priscoficus johnfoxi[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Priscoficus nodulina[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Proceraedis[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species P. imbricataria and P. tegumenis.

Prososthenia esuegirottii[137]

Sp. nov

Valid

Dominici & Forli

Pliocene (Zanclean)

 Italy

A member of the family Hydrobiidae.

Protoplex? zsoldosi[103]

Sp. nov

Valid

Vicián & Kovács

Eocene (Lutetian)

Csernye Formation

 Hungary

A member of the family Cymatiidae.

Proviviparus[138]

Gen. et sp. nov

Valid

Frese & Ponder

Late Jurassic

Talbragar Fish Bed

 Australia

A member of the family Viviparidae. Genus includes new species P. talbragarensis

Pseudoananias[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" subgranulata Münster (1841).

Pseudokatosira uhlhingleyorum[139]

Sp. nov

Valid

Gründel, Davies & Simonsen

Early Jurassic (Pliensbachian)

Pseudolatirus acuelatus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Pseudolatirus.

Pseudolatirus boettgeri[140]

Nom. nov

Valid

Kovács

Miocene

 Romania

A species of Pseudolatirus; a replacement name for Fusus sublamellosus Boettger (1906).

Pseudosassia[103]

Gen. et comb. et 2 sp. nov

Valid

Vicián & Kovács

Paleocene–Miocene

 Austria
 Belgium
 Bulgaria
 Czech Republic
 Denmark
 France
 Germany
 Hungary
 Italy
 Netherlands
 Poland
 Romania
 Serbia
 Slovakia
 Turkey
 Ukraine
 United Kingdom

A member of the family Cymatiidae. The type species is "Triton" flandricum Koninck (1838); genus also includes new species P. gurdoni and P. traceyi.

Pseudoterebellum[141]

Gen. et comb. nov

Valid

Maxwell, Rymer & Congdon

Eocene

Ghazij Shales

 Indonesia
 Pakistan
 Saudi Arabia

A member of the family Seraphsidae. The type species is "Terebellum" diversiornatum Eames (1952); genus also includes "Terebellum" saudiensis Abbass (1972) and "Terebellum" olympiae Rolando (2001).

Ptychatractus cataractus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Ptychatractus.

Pulchraspira[113]

Gen. et sp. nov

In press

Yu, Salvador & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Genus includes new species P. teneristoma.

Puposyrnola? confusa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Possibly a species of Puposyrnola.

Puposyrnola multibaca[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Puposyrnola.

Puposyrnola obliqua[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Puposyrnola.

Puposyrnola shirleyae[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Puposyrnola.

Pusia aturensis[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Pusia.

Pusia confunda[84]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

A species of Pusia.

Pusia? falsitranssylvanica[84]

Nom. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

 Romania

Possibly a species of Pusia; a replacement name for Mitra (Costellaria) recticosta var. transsylvanica Boettger (1902).

Pusia subpyrenaica[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Pusia.

Pusia virodunensis[91]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Pusia.

Pustularia korneli[133]

Sp. nov

Valid

Fehse & Vicián

Miocene

 Indonesia

A species of Pustularia.

Pyrgiscus elegans[142]

Sp. nov

Valid

Tabanelli et al.

Pliocene

 Italy

A species of Pyrgiscus.

Pyrgiscus jaapmulderi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Pyrgiscus.

Pyrgulina marliesae[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Pyrgulina.

Retusa (Cylichnina) praemoryi[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Retusa.

Rhaphaulus zhuoi[113]

Sp. nov

In press

Yu, Salvador & Jarzembowski

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

Rinaldoella tornata[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Ringicula taenia[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

A species of Ringicula.

Sansonia texana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Sassella[73]

Gen. et 2 sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species S. mixta and S. frosti.

Scalptia nemethi[143]

Sp. nov

Valid

Kovács & Vicián

Miocene

Lajta Formation

 Hungary

A species of Scalptia.

Scaphella kendrae[144]

Sp. nov

Valid

Petuch & Berschauer

Pliocene (Zanclean)

Murdock Formation

 United States
( Florida)

A species of Scaphella.

Schizogonium undae[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Seraphs kaindraperi[141]

Sp. nov

Valid

Maxwell, Rymer & Congdon

Eocene (Priabonian)

 Jamaica

A member of the family Seraphsidae.

Similalvania[76]

Gen. et comb. nov

Valid

Chirli & Forli

Pliocene

 Italy

A member of the family Rissoidae. The type species is "Alvania" magistra Chirli (2006)

Solariella microstriata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Solariella.

Solariorbis planulatis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Squamoworthenia[127]

Gen. et comb. nov

Valid

Mazaev

Carboniferous

 Australia
 Germany
 Kazakhstan

Genus includes S. duponti (Holzapfel, 1889) and "Worthenia" crenilunula Yoo (1994).

Stellaria striatissima[75]

Sp. nov

Valid

Raven

Miocene

 Malaysia

A species of Stellaria.

Striacingulum[74]

Gen. et comb. nov

Valid

Karapunar & Nützel

A member of Pleurotomariida. The type species is "Pleurotomaria" cancellatocingulata Klipstein (1844).

Striaticostatum figus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Stuckenbergispira dombarensis[127]

Sp. nov

Valid

Mazaev

Early Carboniferous

 Kazakhstan

Sveltella novus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Sveltella.

Sveltia sofiae[145]

Sp. nov

Valid

Landau & da Silva

Pliocene

 Portugal

A species of Sveltia.

Syrnola debrae[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

A species of Syrnola.

Taioma? argilla[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Taioma calxa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Taranis occulta[72]

Sp. nov

Valid

Brunetti & Cresti

Pliocene

 Italy

A species of Taranis.

Teinostoma regularis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Teinostoma.

Terebellum humilispirum[75]

Sp. nov

Valid

Raven

Miocene

 Malaysia

A species of Terebellum.

Terebra (Mirula) mexia[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Terebra.

Terebrifusus megahani[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Thylacodes devriesi[146]

Sp. nov

In press

Sanfilippo et al.

Miocene

 Peru

A species of Thylacodes.

Tornatellaea globosa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Tornatellaea linifera[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Tripsycha strapplei[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Tritia miniridibunda[86]

Sp. nov

Valid

Lozouet

Miocene

 France

A species of Tritia.

Tritia praebasteroti[86]

Sp. nov

Valid

Lozouet

Oligocene (Chattian)

 France

A species of Tritia.

Tritiaria formosa[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Trivia disjuncta[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Trivia.

Trivia simplex[136]

Sp. nov

Valid

Fehse

Late Oligocene

 France

A species of Trivia.

Trona colossus[118]

Sp. nov

Valid

Groves & Landau

Early–middle Miocene

Baitoa Formation

 Dominican Republic

A species of Trona.

Truncatella jiaozhouensis[147]

Sp. nov

In press

Yu et al.

Late Cretaceous

Jiaolai Basin

 China

A species of Truncatella.

Tumidosipho[73]

Gen. et sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Genus includes new species T. tehuacana.

Turbo (Marmarostoma) ishidai[148]

Sp. nov

Valid

Tomida, Sano & Kase

Miocene

Ena Limestone

 Japan

A species of Turbo.

Turbonilla bincki[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla bongiardinoi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla crovatoi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla malacitana[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla mauroi[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla plioalboranensis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turbonilla velerinensis[106]

Sp. nov

Valid

Landau & Micali

Pliocene

 Spain

A species of Turbonilla.

Turricula frosti[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Turricula.

Turricula imbricata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Turricula.

"Turritella" calax[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Umbilia furneauxensis[149]

Sp. nov

Valid

Southgate, Militz & Roberts

Pliocene

 Australia

A species of Umbilia.

Unitas duomillias[110]

Sp. nov

Valid

Garvie

Eocene

Cook Mountain Formation

 United States
( Texas)

Unitas euani[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Unitas petiti[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Valvata jiaolaiensis[147]

Sp. nov

In press

Yu et al.

Late Cretaceous

Jiaolai Basin

 China

A species of Valvata.

Vanikoropsis bashforthi[150]

Sp. nov

Valid

Schnetler & Nielsen

Paleocene (Danian)

Agatdal Formation

 Greenland

Possibly a member of the family Vanikoridae.

Vanikoropsis jakobseni[150]

Sp. nov

Valid

Schnetler & Nielsen

Paleocene (Danian)

Agatdal Formation

 Greenland

Possibly a member of the family Vanikoridae.

Vanikoropsis mortenseni[150]

Sp. nov

Valid

Schnetler & Nielsen

Paleocene (Selandian)

Kerteminde Marl

 Denmark

Possibly a member of the family Vanikoridae.

Varicospira bekenuensis[75]

Sp. nov

Valid

Raven

Miocene

 Malaysia

A species of Varicospira.

Varicospira reticulata[75]

Sp. nov

Valid

Raven

Miocene

 Malaysia

A species of Varicospira.

Vermicularia? columella[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Vexillum transalpinum[84]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Hungary
 Romania
 Turkey

A species of Vexillum.

Vitrinella discus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Vitrinella.

Vitrinella praelaevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Vitrinella.

Volutifusus dougsheltoni[144]

Sp. nov

Valid

Petuch & Berschauer

Pliocene (Piacenzian)

Tamiami Formation

 United States
( Florida)

A species of Volutifusus.

Volutifusus kissimmeensis[144]

Sp. nov

Valid

Petuch & Berschauer

Pliocene (Piacenzian)

Tamiami Formation

 United States
( Florida)

A species of Volutifusus.

Volutomorpha ultima[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Wormsina[102]

Gen. et comb. nov

Valid

Harzhauser & Landau

Miocene

 Romania

A member of the family Mitridae. The type species is "Mitra (Cylindra)" transsylvanica Hoernes & Auinger (1880).

Wortheniella klipsteini[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Wortheniella paolofedelei[74]

Sp. nov

Valid

Karapunar & Nützel

Late Triassic

San Cassiano Formation

 Italy

A member of Pleurotomariida.

Zafrona sphaerocorrugata[89]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

 Austria
 Bosnia and Herzegovina
 Bulgaria
 Hungary
 Romania
 Ukraine

A species of Zafrona.

Zebinella tehuacana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Zebinella.

Zonaria vokesae[118]

Sp. nov

Valid

Groves & Landau

Early–middle Miocene

Baitoa Formation

 Dominican Republic

A species of Zonaria.

Bivalves[edit]

New taxa[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Acar beadata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Acar.

Amphidonte mirkamalovi[151]

Sp. nov

Valid

Metelkin & Kosenko

Early Cretaceous (Aptian)

An oyster.

Arcopsis subrhombus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Arcopsis tumulus[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Arkhanguelskya[152]

Gen. et comb. nov

Valid

Cooper & Leanza

Cretaceous

A member of the family Buchotrigoniidae. Genus includes "Trigonia" weberi Arkhangelsky (1916).

Baluchicardia? eoplata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Bengtsonigonia[153]

Nom. nov

Valid

Ceccolini & Cianferoni

Early Cretaceous

Riachuelo Formation

 Brazil

A replacement name for Bengtsonella Cooper & Leanza (2019).

Bohemigonia[154]

Gen. et sp. et comb. nov

Valid

Cooper

 Czech Republic

A member of the family Megatrigoniidae. The type species is B. frici; genus also includes "Trigonia" debuchii Geinitz (1840).

Caryocorbula? praefigera[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Chama ferrata[155]

Sp. nov

Valid

Berezovsky

Eocene

 Ukraine

A species of Chama.

Chama subcalcarata[155]

Sp. nov

Valid

Berezovsky

Eocene

 Ukraine

A species of Chama.

Corbulamella palaeocenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Crassatellina minuta[156]

Sp. nov

Valid

Jaitly et al.

Late Cretaceous (Turonian)

Nodular Limestone Formation

 India

A member of the family Lucinidae belonging to the subfamily Eriphylinae.

Crassatellina paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Cuspidaria texana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Cuspidaria.

Diplodonta frosti[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Diplodonta paleocaenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Dorbigonia[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

 France

A member of the family Megatrigoniidae. Genus includes "Trigonia" disparilis d'Orbigny (1844).

Elianegonia[153]

Nom. nov

Valid

Ceccolini & Cianferoni

Late Cretaceous

 Madagascar

A replacement name for Elianella Cooper (2015).

Eomiltha strangensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Freneixicardia picturata[157]

Sp. nov

Valid

Berezovsky

Eocene

 Ukraine

A cockle.

Frenguelliella eopacifica[158]

Sp. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic

 Argentina

A member of Trigoniida belonging to the family Frenguelliellidae.

Glycymeris laevigata[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Glycymeris.

Grammatodon superstes[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Gregariella frosti[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Gregariella.

Himenourella[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

 Japan
 Russia

A member of the family Megatrigoniidae. Genus includes "Trigonia" jimbonis Strand (1928), "Trigonia" sawatai Yehara (1923), "Apiotrigonia" subjimboi Miroljubov in Vereshchagin et al. (1965) and "Trigonia" subovalis Jimbo (1894).

Kolymia rutskovi[159]

Sp. nov

Valid

Biakov in Biakov et al.

Permian

 Russia

Kushmurunskya[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

 Kazakhstan

A member of the family Megatrigoniidae. Genus includes "Trigonia" gladkowskii Shcheglova-Borodina (1960).

Kyzylkumgonia[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

A member of the family Megatrigoniidae. Genus includes "Trigonia" syrdariensis Arkhangelsky (1916), "Trigonia" turcmenensis Arkhangelsky (1916) and "Megatrigonia" tagamensis Belyakova (1980).

Loxocardium marmoreum[157]

Sp. nov

Valid

Berezovsky

Eocene

 Ukraine

A cockle.

Lucina crassiplica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lucina.

Lucina multivittae[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lucina.

Lucina tehuacanensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Lucina.

Maedagonia[154]

Gen. et comb. et nom. nov

Valid

Cooper

Cretaceous

 Japan
 Russia

A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" mikasaensis Tashiro (1979), "Apiotrigonia" ashizawaensis Saito (1962), "Apiotrigonia" crassoradiata Nakano (1957), "Apiotrigonia" futabaensis Maeda & Kawabe (1967), "Apiotrigonia" hetonaiana Tashiro (1978), "Apiotrigonia" hironoensis Maeda & Kawabe (1967), "Apiotrigonia" jimboi Nakano (1957), "Apiotrigonia" obliquecostata Nakano (1957), "Apiotrigonia" obsoleta Nakano (1957), "Apiotrigonia" orikiensis Maeda & Kawabe (1967), "Apiotrigonia" undulosa Nakano (1957) and "Apiotrigonia" utoensis Tashiro (1972), as well as M. yabei (a replacement name for Trigonia subovalis var. minor Yabe & Nagao, 1925).

Martesia mazanticus[160]

Sp. nov

Valid

Castañeda-Posadas, Trujillo-Hernández & Zúñiga-Mijangos

Miocene

Mexican amber

 Mexico

A species of Martesia.

Matsudagonia[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

 Japan

A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" dubia Tashiro (1979).

Moerickella[158]

Gen. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic

 Argentina

A member of Trigoniida, most likely the oldest member of the family Myophorellidae.

Naiadites devonicus[161]

Sp. nov

Valid

Scholze & Gess

Devonian (Famennian)

Witpoort Formation

 South Africa

Opis reticulata[156]

Sp. nov

Valid

Jaitly et al.

Late Cretaceous (Turonian)

Nodular Limestone Formation

 India

A member of the family Lucinidae belonging to the subfamily Opinae.

Ostrea jibananandai[162]

Sp. nov

Valid

Halder & Mitra

Eocene (Ypresian)

Cambay Shale

 India

A species of Ostrea.

Palaeolignopholas[163]

Gen. et sp. nov

Valid

Bolotov et al.

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of the family Pholadidae belonging to the subfamily Martesiinae. The type species is P. kachinensis.

Pandora? paleocenica[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Parmicorbula? sublaevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Perezigonia[153]

Nom. nov

Valid

Ceccolini & Cianferoni

Middle Jurassic

 Chile

A replacement name for Perezella Cooper (2015).

Pojetaia robsonae[164]

Sp. nov

Valid

Peel & Skovsted

Cambrian Stage 4

Kane Basin Formation

 Canada
( Nunavut)

Porterius? strangensis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Poultoniella[154]

Gen. et comb. nov

Valid

Cooper

Cretaceous

 Canada
( British Columbia)

A member of the family Megatrigoniidae. Genus includes "Trigonia" newcombei Packard (1921).

Promyophorella basoaltorum[158]

Sp. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic

 Argentina

A member of Trigoniida belonging to the family Myophorellidae.

Promyophorella? sanjuanina[158]

Sp. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic

 Argentina

A member of Trigoniida belonging to the family Myophorellidae.

Pseudapiotrigonia[154]

Gen. et comb. nov

Valid

Cooper

 United States
( South Dakota
 Utah)

A member of the family Vaugoniidae. Genus includes "Trigonia" sturgisensis Whitfield & Hovey (1906), as well as "Vaugonia" utahensis Imlay (1964).

Pseudovaugonia[158]

Gen. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic

 Argentina

A member of Trigoniida belonging to the family Myophorellidae.

Pteromeris nodulina[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Pteromeris.

Repmanigonia[153]

Nom. nov

Valid

Ceccolini & Cianferoni

Jurassic

 Russia

A replacement name for Repmania Kolesnikov (1980).

Schedocardia imperfecta[157]

Sp. nov

Valid

Berezovsky

Eocene

 Ukraine

A cockle.

Sinoliaoningia[153]

Nom. nov

Valid

Ceccolini & Cianferoni

Late Triassic

Yangcaogou Formation

 China

A replacement name for Liaoningia Yu & Dong (1993).

Sokolovskya[154]

Gen. et comb. nov

Valid

Cooper

A member of the family Megatrigoniidae. Genus includes "Apiotrigonia" levinae Pojarkova (1990) and "Apiotrigonia" khoresmensis Belyakova (1965).

Sonoragonia[154]

Gen. et comb. et nom. nov

Valid

Cooper

Early Cretaceous

Lowell Formation

 United States
( Arizona)

A member of the family Megatrigoniidae. Genus includes "Trigonia" cragini Stoyanow (1949), as well as S. scotti (a replacement name for Trigonia kitchini Stoyanow, 1949).

Sphenotomorpha orientalis[165]

Sp. nov

Valid

Dalenz Farjat

Devonian (Pragian)

Icla Formation

 Bolivia

A member of Modiomorphida belonging to the family Modiomorphidae.

Sunetta crassatelliformis[166]

Sp. nov

Valid

Haga & Fukuda in Fukuda et al.

Early Pleistocene

 Japan

A species of Sunetta.

Sunetta nomurai[166]

Sp. nov

Valid

Haga & Fukuda in Fukuda et al.

Pleistocene

 Taiwan

A species of Sunetta.

Syndesmyella pliominor[167]

Sp. nov

Valid

Brunetti & Cresti

Early Pliocene

 Spain

A member of the family Semelidae.

Tellina brevis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A species of Tellina.

Terrypoultoniella[158][168]

Gen. et sp. nov

Valid

Echevarría, Damborenea & Manceñido

Early Jurassic (PliensbachianToarcian)

 Argentina
 Portugal?

A member of Trigoniida belonging to the family Frenguelliellidae. The type species is "Poultoniella" jaworskii Echevarría, Damborenea & Manceñido (2021); genus might also include "Trigonia" malladae Choffat (1885).

Tindaria hamuroi[169]

Sp. nov

Valid

Amano & Haga

Miocene

Higashibessho Formation

 Japan

A member of the family Tindariidae.

Titomaya[170]

Gen. et comb. et sp. nov

Valid

Alvarez & del Río

Paleocene (Danian)

Lefipán Formation
Salamanca Formation

 Argentina

A member of Venerida belonging to the group Veneroidea. Genus includes "Meretrix" chalcedonica, as well as new species T. longobucca.

Tivelina texana[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Tivelina trigonalis[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

Unionites kobyumensis[171]

Sp. nov

Valid

Biakov in Biakov & Kutygin

Late Permian

 Russia

Vaccinites alencasteri[172]

Sp. nov

Valid

Mitchell

Late Cretaceous (probably late Turonian or early Coniacian)

Cuautla Formation

 Mexico

A rudist bivalve.

Vaccinites temazcali[172]

Sp. nov

Valid

Mitchell

Late Cretaceous (Campanian)

Temazcal Limestone

 Mexico

A rudist bivalve.

Villamilia[152]

Gen. et comb. nov

Valid

Cooper & Leanza

Cretaceous

 Colombia

A member of the family Buchotrigoniidae. Genus includes "Buchotrigonia" etayoi Villamil (1992), "Trigonia" humboldtii Buch (1839), "Buchotrigonia" kauffmani Villamil (1992), "Buchotrigonia" perezi Villamil (1992) and "Vaugonia niranohamensis" santamariae Geyer (1973; raised to the rank of the species V. santamariae).

Weyla santuccii[173]

Sp. nov

Valid

Damborenea et al.

Early Jurassic (Hettangian)

Pogibshi Formation

 United States
( Alaska)

A scallop.

Whitfieldiella luceae[172]

Sp. nov

Valid

Mitchell

Late Cretaceous (Campanian)

Clifton Limestone
Cotui Limestone

 Jamaica
 United States
( Puerto Rico)

A rudist bivalve.

Yunannia[174]

Gen. et 2 sp. nov

Valid

Zhang & Niu in Zhang et al.

Ordovician

Dongchong Formation

 China

A member of Pteriomorphia of uncertain phylogenetic placement. The type species is Y. gankengensis; genus also includes Y. yunkaiensis.

Research[edit]

  • A study on the evolution and possible causes of the extinction of alatoconchid bivalves is published by Chen et al. (2021).[175]
  • A study on the arcoid bivalve diversity during the Eocene doubthouse interval of global climate cooling is published by Hickman (2021).[176]

Other molluscs[edit]

Name Novelty Status Authors Age Type locality Country Notes Images

Anabarella? navaranae[177]

Sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

Cadulus praepalmerae[73]

Sp. nov

Valid

Garvie

Paleocene (Danian)

Kincaid Formation

 United States
( Texas)

A tusk shell, a species of Cadulus.

Catalanispira prima[105]

Sp. nov

Valid

Ebbestad & Cope

Ordovician (Floian)

 United Kingdom

A member of the family Onychochilidae.

Figurina? polaris[177]

Sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

A helcionellid. Subsequently made the type species of a separate genus Sermeqiconus.[178]

Hippopharangites groenlandicus[177]

Sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

A member of the stem group of Aculifera.

Holoornatus[179]

Gen. et sp. nov

In press

Zhang et al.

Cambrian (Fortunian)

 China

A maikhanellid. Genus includes new species H. strigatus.

Inughuitoconus[177]

Gen. et sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

A member of the stem group of Aculifera. Genus includes new species I. borealis.

Lepidozona luzanovkensis[180]

Sp. nov

Valid

Sirenko

Paleocene

Luzanovka Beds

 Ukraine

A chiton, a species of Lepidozona.

Minodentalium[181]

Gen. et 2 sp. et comb. nov

Valid

Asato & Kase

Permian

Akasaka Limestone

 Belgium
 Japan
 New Zealand
 United States

A tusk shell. Genus includes new species M. hayasakai and M. okumurai, as well as "Plagioglypta" furcata Waterhouse (1980), "Pl." girtyi Knight (1940), "Pl." subannulata Easton (1962), "Dentalium" ingens De Koninck (1843), "D." meekianum Geinitz (1866), "Pl." prosseri Morningstar (1922), "Dentalium" priscum Münster in Goldfuss (1842) and "D." herculeum De Koninck (1863).

Ocruranus? kangerluk[177]

Sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

A member of the stem group of Aculifera.

Prodentalium onoi[181]

Sp. nov

Valid

Asato & Kase

Permian

Akasaka Limestone

 Japan

A tusk shell.

Progadilinoides[182]

Gen. et sp. nov

Valid

Guzhov

Early Cretaceous (Aptian)

 Russia

A tusk shell belonging to the family Pulsellidae. The type species is P. balakovensis.

Pseudomyona groenlandica[183]

Sp. nov

Valid

Peel

Cambrian (Drumian)

Fimbuldal Formation

 Greenland

A member of Rostroconchia belonging to the group Tuarangiida and the family Tuarangiidae.

Qingjianglepas[184]

Gen. et sp. nov

In press

Li et al.

Cambrian

 China

A minute limpet-shaped shell, probably belonging to a helcionelloid. Genus includes new species Q. elegans.

Stenotheca? higginsi[177]

Sp. nov

Valid

Peel

Cambrian Stage 4

Aftenstjernesø Formation

 Greenland

General research[edit]

  • A study on the relationship between temperature and diversity in the fossil record of marine molluscs over the last 145 million years is published by Boag, Gearty & Stockey (2021).[185]
  • A study on the relationship between the species richness of shallow-marine molluscs, the number of ecological functional groups, and oceanic temperature in the fossil record of New Zealand over the last ~40 million years is published by Womack et al. (2021).[186]

References[edit]

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  2. ^ a b Korn, D.; Ghaderi, A.; Devaere, L.; Khanehbad, M. (2021). "Ammonoids from the Carboniferous-Permian boundary of east-central Iran". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (3): 335–345. doi:10.1127/njgpa/2021/1015. S2CID 239690567.
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  23. ^ a b c Matsukawa, M. (2021). "Aptian (Lower Cretaceous) ammonite fauna of the Todai Formation, Nagano Prefecture, Japan". Cretaceous Research. 126: Article 104771. Bibcode:2021CrRes.12604771M. doi:10.1016/j.cretres.2021.104771.
  24. ^ Polkovoy, K. S. (2021). "A new genus Kislovodskoceras (Desmoceratidae, Ammonoidea) from the middle Aptian of the vicinity of Kislovodsk (northern Caucasus)". Paleontological Journal. 55 (5): 520–528. doi:10.1134/S0031030121050129. S2CID 238640207.
  25. ^ Goy, A.; Comas-Rengifo, M. J. (2021). "Nejdia (Hildoceratidae, Ammonoidea) in the Toarcian (Lower Jurassic) of the Iberian Peninsula. Systematics, phylogenetic affinities and palaeobiogeography". Historical Biology. 33 (12): 3173–3190. doi:10.1080/08912963.2020.1853719. S2CID 233815255.
  26. ^ a b c Jenks, J. F.; Maekawa, T.; Ware, D.; Shigeta, Y.; Brayard, A.; Bylund, K. G. (2021). "Late Griesbachian (Early Triassic) ammonoids and nautiloids from the Dinwoody Formation at Crittenden Springs, Elko County, Nevada". New Mexico Museum of Natural History and Science Bulletin. 86: 1–23.
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