Gomphonema

Gomphonema
Diatom Gomphonema
Scientific classification
Domain:	Eukaryota
Clade:	Diaphoretickes
Clade:	SAR
Clade:	Stramenopiles
Phylum:	Gyrista
Subphylum:	Ochrophytina
Class:	Bacillariophyceae
Order:	Cymbellales
Family:	Gomphonemataceae
Genus:	Gomphonema C.G.Ehrenberg, 1832

Gomphonema is a genus of diatoms belonging to the family Gomphonemataceae.

The genus has cosmopolitan distribution.

Species

As accepted by WoRMS;

• Gomphonema acceptatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema acuminatum Ehrenberg, 1832
• Gomphonema adhikarii C.Radhakrishnan, Sudipta K.Das, Kociolek et B.Karthick in Radhakrishnan et al., 2020
• Gomphonema affine Kützing, 1844
• Gomphonema africanum G.S.West, 1907
• Gomphonema alavariense C. Delgado, S. Blanco, L. Gonzalez-Paz & S.F.P. Almeida in Gonzalez-Paz et al., 2020
• Gomphonema angkoricum Cejudo-Figueiras, S.Blanco & Álvarez-Blanco, 2012
• Gomphonema angusticapitatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema angusticlavatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema angustissimum F.Hustedt
• Gomphonema angustum C.Agardh, 1831
• Gomphonema anjae Lange-Bertalot & Reichardt
• Gomphonema asiaticum Liu & Kociolek, 2013
• Gomphonema augur Ehrenberg, 1840
• Gomphonema auguriforme Levkov, Mitic-Kopanja, Wetzel & Ector in Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema auritum A.Braun ex Kützing, 1849
• Gomphonema balatonis Pantocsek
• Gomphonema berggreni
• Gomphonema berggrenii Cleve
• Gomphonema bergii Skvortzov & Meyer
• Gomphonema bicepiformis Wei Zhang & Kociolek, 2018
• Gomphonema bigutianchnensis Y.Li in Liao & Y.Li, 2018
• Gomphonema bohemicum Reichelt & Fricke, 1902
• Gomphonema boreale E.V.Østrup
• Gomphonema brasiliensoide Metzeltin, Lange-Bertalot & García-Rodríguez, 2005
• Gomphonema bukycanyona L.N.Bukhtiyarova, 2013
• Gomphonema californicum Stancheva & Kociolek, 2016
• Gomphonema candelariae J.Frenguelli
• Gomphonema cantalicum J.-J.Brun & Héribaud-Joseph
• Gomphonema capitatum Ehrenberg, 1838
• Gomphonema capitoccultum Reichardt
• Gomphonema capsulare J.A.Ress, E.W.Thomas & Kociolek, 2016
• Gomphonema carolinense Hagelstein, 1939
• Gomphonema catarinensis G.Krasske, 1948
• Gomphonema changyangicum Li, Shi & Lei, 1999
• Gomphonema charcotii M.Peragallo
• Gomphonema chemeron Jeff.R.Stone, Westover & Kociolek
• Gomphonema chinense Liu & Kociolek, 2013
• Gomphonema clavatoides H.P.Gandhi, 1960
• Gomphonema clavatuloides Rimet, D.G. Mann, Trobajo et N. Abarca in Rimet et al., 2018
• Gomphonema clavatum Ehrenberg, 1832
• Gomphonema clerici Frenguelli, 1923
• Gomphonema clevei Fricke, 1902
• Gomphonema confusum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema correntinum Frenguelli, 1933
• Gomphonema dalatica Glushchenko, Kulikovskiy & Kociolek, 2017
• Gomphonema darwinii L.L.Bahls, 2014
• Gomphonema delicatula Skvortzov, 1937
• Gomphonema dharwarensis H.P.Gandhi, 1956
• Gomphonema dichotiforme Z.X.Shi, 2004
• Gomphonema difformum B.Karthick & J.P.Kociolek, 2011
• Gomphonema diminutum B.Karthick & J.P.Kociolek, 2011
• Gomphonema dirangense S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
• Gomphonema distincte-striatum Kociolek, J. P. & Woodward, J.
• Gomphonema dojranense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema doonense B.Karthick, R.Nautiyal & Kociolek, 2015
• Gomphonema dubravicense Pantocsek
• Gomphonema eileencoxiae (Bahls, 2017) Bahls, C.E.Wetzel & Ector, 2018
• Gomphonema elegantissimum E.Reichardt & H.Lange-Bertalot, 2011
• Gomphonema elongatum W.Smith, 1855
• Gomphonema eriensioides Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
• Gomphonema evolutionense L.L.Bahls, 2014
• Gomphonema frequentiformis (Metzeltin & Krammer) C.E.Wetzel & F.F.Almeida in Almeida et al., 2020
• Gomphonema frickei O.Müller, 1905
• Gomphonema gallaudi Héribaud-Joseph
• Gomphonema gandhii B.Karthick & J.P.Kociolek, 2011
• Gomphonema gautieriforme D.Mitic-Kopanja, C.E.Wetzel, L.Ector & Z. Levkov, 2014
• Gomphonema genestoermeri Liu & Kociolek, 2013
• Gomphonema graciledictum E.Reichardt, 2015
• Gomphonema grande B. Karthick, Kociolek, J.C. Taylor & Cocquyt, 2016
• Gomphonema gregarium J.P Kociolek & Jeff Woodward, 2016
• Gomphonema hedinii Hustedt, 1922
• Gomphonema heideni F.Fricke
• Gomphonema helveticum Brun, 1895
• Gomphonema himalayaense Jüttner in Jüttner et al., 2018
• Gomphonema hinziae Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema hristovskii Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema hubeicum Li, Shi & Lei, 1999
• Gomphonema idsbense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema indistinctum Kociolek, J. P. & Woodward, J.
• Gomphonema innata Skvortzov, 1937
• Gomphonema insigne Gregory, 1856
• Gomphonema insuetum Kociolek in Kociolek et al., 2014
• Gomphonema insularum Kociolek, J. P. & Woodward, J.
• Gomphonema interruptum J.YChen & H.Z.Zhu, 1994
• Gomphonema intricatoides You & Kociolek, 2015
• Gomphonema intricatum Kützing, 1844
• Gomphonema italicum Kützing, 1844
• Gomphonema jablanicense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema jahniana Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
• Gomphonema johnsonii L.Bahls, 2013
• Gomphonema juettnerii B.Karthick, R.Nautiyal & Kociolek, 2015
• Gomphonema juriljii Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema kalahariense Jeff.R.Stone & Kociolek
• Gomphonema kallarense Pardhi et al., 2020
• Gomphonema kaznakowi K.S.Mereschkowsky
• Gomphonema kerguelenensis E.E.Manguin, 1954
• Gomphonema kezlyae Pardhi et al., 2020
• Gomphonema kociolekii R.Jahn & N.Abarca
• Gomphonema kozufense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema lacifrigidum M.Kulikovskiy, J.P.Kociolek & Solak in Kociolek & Guiry, 2018
• Gomphonema lacusrankala Gandhi, 1958
• Gomphonema lacusrankaloides B.Karthick & J.P.Kociolek, 2012
• Gomphonema lacus-victoriense E.Reichardt, 2007
• Gomphonema lagerheimei A.Cleve, 1895
• Gomphonema lagerheimii A.Cleve, 1895
• Gomphonema lapponicum (A.Cleve) Cleve-Euler
• Gomphonema latelanceolatum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema lateripunctatum Reichardt & Lange-Bertalot, 1991
• Gomphonema liebscheri E.Reichardt, 2012
• Gomphonema longiceps Ehrenberg, 1854
• Gomphonema lujanensis E.Reichardt & N.Maidana, 2008
• Gomphonema lychnidum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema magniaqua M.Kulikovskiy, J.P.Kociolek & Solak in Kociolek & Guiry, 2018
• Gomphonema magnifica H.P.Gandhi
• Gomphonema mariovense Levkov & Tofilovska in Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema martini F.Fricke
• Gomphonema matanensis D. Kapustin, Kociolek & Kulikovskiy in Kociolek et al., 2018
• Gomphonema mayamae S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
• Gomphonema megalobrebissonii D.A.Chudaev, J.P.Kociolek & M.A.Golobova, 2014
• Gomphonema meridionale J.P.Kociolek & E.W.Thomas, 2009
• Gomphonema metzeltinii You & Kociolek, 2015
• Gomphonema microcapitatum Kulikovskiy, Kociolek & Solak, 2018
• Gomphonema microlanceolatum You & Kociolek, 2015
• Gomphonema microlaticollum Kulikovskiy, Kociolek & Solak, 2018
• Gomphonema micropus Kützing, 1844
• Gomphonema minusculum Krasske
• Gomphonema minutum (C.Agardh) C.Agardh, 1831
• Gomphonema modicum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema moniliforme Gandhi, 1960
• Gomphonema mutunensis Ruwer, Ludwig & Rodrigues, 2019
• Gomphonema nagpurense Sarode & Kamat, 1984
• Gomphonema navicelloides Frenguelli, 1942
• Gomphonema naviculoides W. Smith, 1856
• Gomphonema neotropicum N.Abarca & D.Mora
• Gomphonema nesophilum Kociolek, J. P. & Woodward, J.
• Gomphonema obesoccultum Reichardt
• Gomphonema occidentale
• Gomphonema olivaceum (Lyngbye) Desmazières, 1825
• Gomphonema oregonicum Ehrenberg
• Gomphonema oxycephalum Cleve
• Gomphonema paracapitatum Kulikovskiy, Kociolek & Solak, 2018
• Gomphonema paradaphnoides Cejudo-Figueiras, S.Blanco & Ãlvarez-Blanco, 2012
• Gomphonema paraexilissimum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema parapygmaeum Jüttner & Kociolek in Jüttner et al., 2018
• Gomphonema parasundaense Jüttner in Jüttner et al., 2018
• Gomphonema paratergestinum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema parvicapitatum Kociolek in Kociolek et al., 2014
• Gomphonema parvuliforme Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema parvulum (Kützing) Kützing, 1849
• Gomphonema pelisteriense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema preliciae Levkov, Mitic-Kopanja & E. Reichardt, 2016
• Gomphonema prespanense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema prowsei Glushchenko, Kulikovskiy & Kociolek, 2017
• Gomphonema pseudacuminatum Kulikovskiy, Kociolek & Solak, 2015
• Gomphonema pseudaffine Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema pseudoparvulum Kociolek, J. P. & Woodward, J.
• Gomphonema puiggarianum
• Gomphonema pusillum (Grunow, 1880) Kulikovskiy & Kociolek, 2018
• Gomphonema pygmaeoides You & Kociolek, 2015
• Gomphonema qingyiensis L-X. Zhang, P. Yu & Q-M. You in Zhang et al., 2020
• Gomphonema raraense Jüttner & S.Gurung in Jüttner et al., 2018
• Gomphonema reediae Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema resendei A.R.Moura
• Gomphonema rexlowei Liu & Kociolek, 2013
• Gomphonema rhombicum Fricke, 1904
• Gomphonema rimetiana Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
• Gomphonema ristovskae Levkov & Tofilovska in Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema rostellatum (Kützing) G.L.Rabenhorst
• Gomphonema russicum Kulikovskiy, Kociolek & Solak in Kociolek et al., 2018
• Gomphonema saparense S.K.Das, C.Radhakrishnan, Kociolek & B.Karthick, 2018
• Gomphonema saravathense Gandhi, 1970
• Gomphonema scardicum D.Mitic-Kopanja, C.E.Wetzel, L.Ector & Z.Levkov, 2014
• Gomphonema semiapertum Grunow, 1880
• Gomphonema shanghaiensis Zhang & Kociolek, 2016
• Gomphonema shiwania Y.Liu, Kulikovskiy & Kociolek in Zhang et al., 2020
• Gomphonema sichuanenesis Y.Li & J.P.Kociolek, 2010
• Gomphonema sichuanensis Li & Kociolek in Li et al., 2010
• Gomphonema sierrianum Stancheva & Kociolek, 2016
• Gomphonema sphaerophorum Ehrenberg, 1845
• Gomphonema spicula Gandhi, 1958
• Gomphonema spiculoides H.P.Gandhi
• Gomphonema spirkovskae Levkov & Mitic-Kopanja in Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema stauroneiforme Grunow
• Gomphonema staurophorum (Pantocsek) Cleve-Euler, 1955
• Gomphonema subapicatum F.E.Fritsch & M.F.Rich
• Gomphonema subclavatum (Grunow) Grunow, 1884
• Gomphonema subinsigniforme L.Ge, Y.Liu & J.P.Kociolek, 2014
• Gomphonema sublaticollum Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema submalayense Gandhi, 1970
• Gomphonema subnaviculoides Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema subodhii Jüttner & S.Gurung in Jüttner et al., 2018
• Gomphonema subramosum C.A.Agardh
• Gomphonema subtile Ehrenberg, 1843
• Gomphonema svalbardense E.Pinseel, K.Kopalová & Van der Vijver, 2014
• Gomphonema tamilense B.Karthick & J.P.Kociolek, 2012
• Gomphonema tenoccultum Reichardt
• Gomphonema tenuis Gandhi, 1960
• Gomphonema tergestinum (Grunow in Van Heurck) Fricke, 1902
• Gomphonema truncatum Ehrenberg, 1832
• Gomphonema tumens Kociolek & Stoermer, 1991
• Gomphonema turgidum Ehrenberg, 1854
• Gomphonema undulans I.Ãlvarez-Blanco, C.Cejudo-Figueiras & S.Blanco, 2013
• Gomphonema validum Cleve, 1894
• Gomphonema vardarense Levkov, Mitic-Kopanja & E.Reichardt, 2016
• Gomphonema variabilis A.Jurilj, 1957
• Gomphonema varioreduncum Jüttner, Ector, E. Reichardt, Van de Vijver & E.J.Cox, 2013
• Gomphonema vastum F.Hustedt
• Gomphonema ventricosum Gregory, 1856
• Gomphonema vibrio Ehrenberg, 1843
• Gomphonema vidarbhense Sarode & Kamat, 1984
• Gomphonema williamsii Kociolek & Liu, 2013
• Gomphonema witkowskii Kociolek & Liu, 2013
• Gomphonema wuyiensis W.Zhang & Kociolek in Zhang et al., 2018
• Gomphonema xiantaoicum Z.X.Shi & N.Li in Z.X.Shi, 2014
• Gomphonema xinjiangianum You & Kociolek, 2015
• Gomphonema yangtzensis Li in Li, Gong, Xie & Shen, 2006
• Gomphonema yaominae Li, 2012
• Gomphonema yberiense Frenguelli, 1933
• Gomphonema yunnaniana Y.Liu et Kociolek in Liu et al., 2020

Fossils

Several species are fossils.

• Gomphonema jianghanense Y.L.Li, Z.J.Gong, P.Xie & J.Shen, 2007 †
• Gomphonema jianghanensis Li, Gong, Xie & Shen, 2007 †
• Gomphonema jilinense Huang in Huang et al., 1983 †